Joseph Raymond Coelho 


The Institute for Franciscan Environmental Studies
Biology Program
Quincy University
1800 College Avenue
Quincy, IL 62301 U.S.A.
Phone: 217-228-5432 x3268
FAX: 217-228-5222
Office: North Campus A207


1984. B.S. Biology with honors. University of California, Riverside

1989. Ph.D. Environmental, Population and Organismic Biology. University of Colorado, Boulder.

Professional Organizations


Reprints available by request for most articles.

Coelho, J.R., C.W. Holliday, J.M. Hastings and C.M. Phillips. 2016. Thermal biology of Pacific cicada killers, Sphecius convallis Patton, in the upper Sonoran desert. Journal of Thermal Biology 57:101–109. doi:10.1016/j.jtherbio.2016.03.001

Cooley, J.R., D.C. Marshall, A.F. Richards, R.D. Alexander, M.D. Irwin, J.R. Coelho and C. Simon. 2013. The distribution of periodical cicada Brood III in 1997, with special emphasis on Illinois (Hemiptera: Magicicada spp.). American Entomologist 59:9-14.

Coelho, J.R., J.M. Hastings and C.W. Holliday. 2012. Near-optimal foraging in the Pacific cicada killer Sphecius convallis Patton (Hymenoptera: Crabronidae). Insects 3(1):133-140.

Coelho J.R. 2011. Effects of prey size and load carriage on the evolution of foraging strategies in wasps. In: Predation in the Hymenoptera: An Evolutionary Perspective (Polidori, C., ed.). Transworld Research Network publishing, Kerala, India, p. 23-38.

Coelho, J.R., C.W. Holliday and J.M. Hastings. 2011. The geographic distributions of cicada killers (Sphecius; Hymenoptera, Crabronidae) in the Americas. The Open Entomology Journal 5:31-38.

Coelho, J.R. 2011. Noninsect arthropods in popular music. Insects 2(2):253-263.

Holliday, C.W., J.M. Hastings and J.R. Coelho. 2009. Cicada prey of New World cicada killers, Sphecius spp. (Dahlbom 1843) (Hymenoptera : Crabronidae). Entomological News 120(1):1-17.

Hastings, J.M., C.W. Holliday, and J.R. Coelho. 2008. Body size relationship between Sphecius speciosus (Hymenoptera: Crabronidae) and their prey: prey size determines wasp size. Florida Entomologist 91(4):657-663.

Hastings, J.M., J.R. Coelho, and C.W. Holliday. 2008. Mating at high population density in a colonial territorial wasp, Sphecius speciosus Drury (Hymenoptera : Crabronidae). Journal of the Kansas Entomological Society 81(3):301-308.

Coelho, J.R. and C.W. Holliday. 2008. The effect of hind-tibial spurs on digging rate in female eastern cicada killers. Ecological Entomology 33:403-407.

Coelho, J.R., J.M. Hastings, C.W. Holliday, G. Flure and K. Barnes. 2008. Sexual dimorphism of the femora, tibiae and hind tibial spurs in the eastern cicada killer, Sphecius speciosus Drury (Hymenoptera: Crabronidae) in the United States. Entomological News 119(1):11-18.

Hastings, J.M., P. J. Schultheis, M.K. Whitson, C.W. Holliday, J.R. Coelho, and A.M. Mendell. 2008. DNA barcoding of New World cicada killers (Hymenoptera: Crabronidae). ZooTaxa 1713:27-28.

Coelho, J.R., J.M. Hastings, C.W. Holliday, and A.M. Mendell. 2008. Load carriage during foraging in two species of solitary wasps. Journal of Hymenoptera Research 17(1):57-63.

Coelho, J.R.. 2007. Review of The Sand Wasps: Natural History and Behavior by H.E. Evans and K.M. O’Neill (Harvard, 2007). The Quarterly Review of Biology 82(4):424-425.

Coelho, J.R., C.W. Holliday, J.M. Hastings, E. Maty, M. Swigart and A. Mendell. 2007. Thermoregulation in male western cicada killers (Sphecius grandis Say) in the Chihuahuan desert. Journal of Thermal Biology 32(5):270-275. Abstract

Holliday, C.W. and J.R. Coelho. 2006. Improved key to New World species of Sphecius (Hymenoptera: Crabronidae). Annals of the Entomological Society of America 99(5):793-798.

Coelho, J.R. 2006. Abraham Merritt's The Drone: a forgotten classic of entomological fantasy. American Entomologist 52(2):70-71. [The Drone]

Coelho, J.R. 2004. Insects in rock and roll cover art. American Entomologist 50(3):142-151.

Coelho, J.R. 2003. Thorax temperature and honey-crop load of honey bee undertakers as potential constraints on task performance. Ex Scientia 9:16-25.

Coelho, J.R. 2002. Spurred on to greater depths. Natural History 111(6): 20-22

Coelho, J.R. 2002. Living the Sporting Life: Adventures and Misadventures in the Outdoors., 101 pp.

Coelho, J.R. 2001. The natural history and ecology of antlions (Neuroptera:Myrmeleontidae). Ex Scientia 7:3-12.

Coelho, J.R. and C.W. Holliday. 2001. Effects of size and flight performance on intermale mate competition in the cicada killer, Sphecius speciosus Drury (Hymenoptera: Sphecidae). Journal of Insect Behavior 14(3):345-351. Abstract

Coelho, J.R. 2001. Behavioral and physiological thermoregulation in male cicada killers (Sphecius speciosus) during territorial behavior . Journal of Thermal Biology 26(2):109-116.

Edgar, P.K. and J.R. Coelho. 2000. Load-lifting constraints on provisioning and nest building in the carpenter wasp, Monobia quadridens L. (Hymenoptera: Eumenidae). Journal of Hymenoptera Research 9 (2):370-376.

Coelho, J.R. 2000. Insects in rock and roll music. American Entomologist 46(3):186-200.

Irwin, M.D. and J.R. Coelho. 2000. The distribution of the Iowan Brood of periodical cicadas (Homoptera: Cicadidae: Magicicada spp.) in Illinois. Annals of the Entomological Society of America 93(1):82-89.

Carter, P., J.B. Mitton, T.D. Kocher, and J.R. Coelho. 2000. Maintenance of the ADH polymorphism in tiger salamanders II. Differences in biochemical function among allozymes. Functional Ecology 14(1):70-76. Abstract

Coelho, J.R. and L.D. LaDage. 1999. Foraging capacity of the great golden digger wasp, Sphex ichneumoneus L. Ecological Entomology 24:480-483.

Coelho, J.R. 1999. Body temperature and flight muscle ratio in the burrowing mayfly, Hexagenia bilineata. Journal of Freshwater Ecology 14(3):337-341.

Coelho, J.R. and K. Wiedman.  1999.  Functional morphology of the hind tibial spurs of the cicada killer (Sphecius speciosus Drury).  Journal of Hymenoptera Research 8(1):6-12. 

Coelho, J.R. 1998. An acoustical and physiological analysis of buzzing in cicada killer wasps (Sphecius speciosus). Journal of Comparative Physiology A 183 (6): 745-751. Abstract

Coelho, J.R. 1997. Sexual size dimorphism and flight behavior in cicada killers (Sphecius speciosus). Oikos 79:371-375.

Meehan, R. and Coelho, J.R. 1997. Instructor's manual and test item file to accompany Life 3rd ed by R. Lewis. WCB McGraw Hill, Dubuque, Iowa, 327 pp.

Coelho, J.R. 1996. The flight characteristics of drones in relation to mating. BeeScience 4(1):21-25.

Coelho, J.R. and A.J. Ross. 1996. Body temperature and thermoregulation in two species of yellowjackets, Vespula germanica and V. maculifrons. Journal of Comparative Physiology B: Biochemical, Systemic, and Environmental Physiology 166(1):68-76.

Coelho, J.R., and J. Hoagland. 1995. Load-lifting capacity and foraging of three species of yellowjackets (Vespula) on honey-bee corpses. Functional Ecology 9:171-174.

Ross, A.J., and J.R. Coelho. 1995. The effect of thorax temperature on flight speed in honey bee drones (Apis mellifera). Pages 987-991 in Proceedings of the Undergraduate Research Conference. Volume III (Robert D. Yearout, ed.). University of North Carolina at Asheville, 1149 pp.

Coelho, J.R. and J.B. Sullivan. 1994. Colonization of wildlife nest boxes by honey bee swarms. American Bee Journal 134(10):697-699.

Snyder, G.K., C. Farrelly, and J.R. Coelho. 1992. Capillary perfusion in skeletal muscle. American Journal of Physiology: Heart and Circulatory Physiology 262(31): H828-H832.

Snyder, G.K., C. Farrelly, and J.R. Coelho. 1992. Adaptations in skeletal muscle capillarity following changes in oxygen supply and changes in oxygen demands. European Journal of Applied Physiology and Occupational Physiology 65:158-163.

Coelho, J.R. 1991. The effect of thorax temperature on force production during tethered flight in honeybee (Apis mellifera) drones, workers, and queens. Physiological Zoology 64(3):823-835.

Coelho, J.R. 1991. Heat transfer and body temperature in honey bee drones and workers (Hymenoptera: Apidae). Environmental Entomology 20(6):1627-1635.

Snyder, G.K., J.R. Coelho, and D.R. Jensen. 1991. Body temperature regulation and oxygen consumption in young chicks fed thyroid hormone. Canadian Journal of Zoology 69(7):1842-1847.

Snyder, G.K. and J.R. Coelho. 1989. Microvascular development in chick anterior latissimus dorsi following hypertrophy. Journal of Anatomy 162:215-224.

Coelho, J.R. and A.J. Moore. 1989. Allometry of resting metabolic rate in cockroaches. Comparative Biochemistry and Physiology 94A(4):587-590.

Coelho, J.R. and J.B. Mitton. 1988. Oxygen consumption during hovering is associated with genetic variation of enzymes in honeybees. Functional Ecology 2:141-146.


2009. Illinois Rain Garden Initiative. $500.

Recent Presentations

2011. Peoria Camera Club. "Ghost Town: Ruby, Arizona."

2011. Missouri Nature and Environmental Photographers, St. Louis. "Photographing the Galapagos."

2006. Sigma Zeta National Conference, McKendree College. "Sex, bugs and rock & roll" (keynote address).

2006. Missouri Academy of Science, Kirksville, MO. "Missouri Stream Team as a teaching tool."

2005. Entomological Society of America: "Insects in rock & roll cover art: new trends.”

2005. Missouri Academy of Science, Jefferson City, MO. “Distribution of Cicada Killers (Sphecius) in North America: a First Look.”

2004. Missouri Academy of Science, Kansas City, MO. "Determining the Distribution of a Brood of Periodical Cicadas Using GPS: Methods Used, Lessons Learned." presented in the special symposium Geospatial Technology Applications in Conservation.

Professional Experience

2008 - present. Associate Professor. Institute for Franciscan Environmental Studies, Biology Program, Quincy University, Quincy, IL
2004- 2008. Assistant Professor. Biology Program. Quincy University, Quincy, IL
2001 - 2004. Associate Professor and Chair. Dept. of Biology, Culver-Stockton College.
1999 - 2001. Assistant Professor. Dept. of Biology, Culver-Stockton College, Canton, MO.
1993 - 1999. Assistant Professor. Dept. of Biological Sciences, Western Illinois University, Macomb, IL
1990 - 1993. Assistant Professor. Dept. of Biology, Marian College, Indianapolis, IN.
1990. Postdoctoral Research Associate. Dept. of EPO Biology, University of Colorado, Boulder

[05/28/2011 jrc]